Let's say that you have Skinshifter in play and your opponent attacks with a 3/3 token. You choose to block with Skinshifter and in response to the block, transform him into a 4/4 rhino creature with trample. Then your opponent (in response to you transforming Skinshifter) plays Magma Jet and targets the now transformed Skinshifter. Can you transform Skinshifter a second time provided you have the mana?
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No. the last line of the ability literally states: "Active this ability only once each turn"
Actually, with Rings of Brighthearth, you can manage to transform Skinshifter twice in one turn. (But do note that you can't change modes on the copied ability.)