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Let's say that you have Skinshifter in play and your opponent attacks with a 3/3 token. You choose to block with Skinshifter and in response to the block, transform him into a 4/4 rhino creature with trample. Then your opponent (in response to you transforming Skinshifter) plays Magma Jet and targets the now transformed Skinshifter. Can you transform Skinshifter a second time provided you have the mana?

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With regard to "targets the now transformed Skinshifter" - You seem to have some confusion about what it means to "respond to" something. When your opponent responds to Skinshifter's activated ability, it has not transformed yet. It dies to Magma Jet, and then the ability resolves, but Skinshifter is dead now, so nothing happens. –  Rainbolt May 20 at 16:11

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No. the last line of the ability literally states: "Active this ability only once each turn"

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Actually, with Rings of Brighthearth, you can manage to transform Skinshifter twice in one turn. (But do note that you can't change modes on the copied ability.)

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Whilst that's technically true, it won't help the asker do what they're trying to do. –  Jonathan Hobbs May 21 at 6:47

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